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{{!}} inserts a vertical bar "|" within a template parameter without delimiting the end of that parameter. It should be used only for inserting tables, or table columns, within a template. It is necessary if the character is used in wiki table syntax, or if the parameter is automatically wikilinked and you wish to use a pipe to specify different link text from the target page name. In all other cases, such as to merely display a vertical bar, it's better to use &#124; or the almost equivalent (but verbose) <nowiki>|</nowiki>. See also Wikipedia's Template:!.